In 1556, Don Aguirre leads Spanish conquistadors to South America to search for the Tears of the Moon, a mythical tree whose flowers cure illness, heal injuries, and lift curses. After most conquistadors die, the Puka Michuna tribe heals the survivors with the Tree's flowers. The tribal chief refuses to reveal the Tree's location, so Aguirre stabs him and burns the village. The dying chief curses the conquistadors, making them immortal and unable to leave sight of the Amazon River; the jungle recaptures anyone attempting to escape.
In 1916 London, Dr. Lily Houghton's Tears of the Moon research is presented by her brother, MacGregor, to the Royal Society, claiming that the Tree's flowers could revolutionize medicine and aid the British war effort. The Houghtons request access to a recently acquired arrowhead artifact. Lily believes it and an old Amazon map are the key to finding the Tree. The society denies the request, believing the Tree is a myth and women scientists are inferior. Lily steals the arrowhead, narrowly evading Prince Joachim, an ambitious German royal also seeking the Tree.
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